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Denaturation temperature (*T*~d~) {#Sec10}

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A ChronOS instrument (CHONDROLAB, Australia) was used to determine the denaturation temperature of each protein. The measurement was performed with a step thermal scan at the rate of 0.5 °C/min from 23 to 95 °C. The ellipticity at 222 nm was continuously monitored. The *T*~d~ was determined by the peak of denaturation observed in the graph of ellipticity as a function of temperature.

Circular dichroism {#Sec11}

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CD spectra were determined in a Jasco J-720 spectrometer with a 0.1 cm path length cell at the indicated pHs. The proteins were adjusted to pHs 7.0, 6.5, 6.